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Ying Tian, [email protected] These authors have contributed equally to this work and share first authorship This article was submitted to Cell Death and Survival, a section of the journal Frontiers in Cell and Developmental Biology Disclaimer All claims expressed in this article are solely those of the authors and do not necessarily represent those of their affiliated organizations, or those of the publisher, the editors and the reviewers

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As intrahepatic cholangiopathy remains unresolved by surgical intervention, patients develop cholangitis (50%), portal hypertension (60%), and progressive fibrosis, leading to liver failure (50%) requiring liver transplantation with lifelong immunosuppressants

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